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Wing coupling (Lepidoptera anatomy) : ウィキペディア英語版 | Wing coupling
Some four-winged insect orders, such as the Lepidoptera, have developed a wide variety of morphological wing coupling mechanisms in the imago which render these taxa as "functionally dipterous" (effectively two-winged) for efficient insect flight. All, but the most basal forms, exhibit this wing coupling. The mechanisms are of three different types - jugal, frenulo-retinacular and amplexiform.〔 Subsec. "Wing coupling". Pp 56-60.〕 ==Jugal wing coupling==
The more primitive groups of moth have an enlarged lobe-like area near the basal posterior margin, i.e. at the base of the forewing, called ''jugum'', that folds under the hindwing in flight.〔
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